Depression-era bank robber John Dillinger's (Johnny Depp) charm and audacity endear him to much of America's downtrodden public, but he's also a thorn in the side of J. Edgar Hoover (Billy Crudup) and the fledgling FBI. Desperate to capture the elusive outlaw, Hoover makes Dillinger his first Public Enemy Number One and assigns his top agent, Melvin Purvis (Christian Bale) the task of bringing him in dead or alive.